DAMASCUS, Syria — Syrian government forces pushed into the ancient town of Palmyra, where Islamic State militants appeared on the verge of collapse Thursday, while in Iraq, a military spokesman announced the start of a long-awaited operation to recapture the IS-held northern city of Mosul.

The extremist group has been losing ground in Syria and Iraq for months under a stepped-up campaign of U.S.-led and Russian airstrikes, as well as ground assaults by multiple forces in each country.

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